[Darwinism (1889) by Alfred Russel Wallace]@TWC D-Link book
Darwinism (1889)

CHAPTER III
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That the amount of variation is large, may be seen by comparing it with the actual length of the head (given below the diagram) which was used as a standard in determining the variation, but which itself seems not to have varied.[19] _Variation among Birds_.
Coming now to the class of Birds, we find much more copious evidence of variation.

This is due partly to the fact that Ornithology has perhaps a larger body of devotees than any other branch of natural history (except entomology); to the moderate size of the majority of birds; and to the circumstance that the form and dimensions of the wings, tail, beak, and feet offer the best generic and specific characters and can all be easily measured and compared.

The most systematic observations on the individual variation of birds have been made by Mr.J.A.Allen, in his remarkable memoir: "On the Mammals and Winter Birds of East Florida, with an examination of certain assumed specific characters in Birds, and a sketch of the Bird Faunae of Eastern North America," published in the _Bulletin of the Museum of Comparative Zoology_ at Harvard College, Cambridge, Massachusetts, in 1871.

In this work exact measurements are given of all the chief external parts of a large number of species of common American birds, from twenty to sixty or more specimens of each species being measured, so that we are able to determine with some precision the nature and extent of the variation that usually occurs.
Mr.Allen says: "The facts of the case show that a variation of from 15 to 20 per cent in general size, and an equal degree of variation in the relative size of different parts, may be ordinarily expected among specimens of the same species and sex, taken at the same locality, while in some cases the variation is even greater than this." He then goes on to show that each part varies to a considerable extent independently of the other parts; so that when the size varies, the proportions of all the parts vary, often to a much greater amount.

The wing and tail, for example, besides varying in length, vary in the proportionate length of each feather, and this causes their outline to vary considerably in shape.


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