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Darwinism (1889)

CHAPTER III
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Thus, Dr.S.P.
Woodward states that many present a most perplexing amount of variation, resulting (as he supposes) from supply of food, variety of depth and of saltness of the water; but we know that many variations are quite independent of such causes, and we will now consider a few cases among the land-mollusca in which they have been more carefully studied.
In the small forest region of Oahu, one of the Sandwich Islands, there have been found about 175 species of land-shells represented by 700 or 800 varieties; and we are told by the Rev.J.T.Gulick, who studied them carefully, that "we frequently find a genus represented in several successive valleys by allied species, sometimes feeding on the same, sometimes on different plants.

In every such case the valleys that are nearest to each other furnish the most nearly allied forms; _and a full set of the varieties of each species presents a minute gradation of forms between the more divergent types found in the more widely separated localities_." In most land-shells there is a considerable amount of variation in colour, markings, size, form, and texture or striation of the surface, even in specimens collected in the same locality.

Thus, a French author has enumerated no less than 198 varieties of the common wood-snail (Helix nemoralis), while of the equally common garden-snail (Helix hortensis) ninety varieties have been described.

Fresh-water shells are also subject to great variation, so that there is much uncertainty as to the number of species; and variations are especially frequent in the Planorbidae, which exhibit many eccentric deviations from the usual form of the species--deviations which must often affect the form of the living animal.

In Mr.Ingersoll's Report on the Recent Mollusca of Colorado many of these extraordinary variations are referred to, and it is stated that a shell (Helisonia trivolvis) abundant in some small ponds and lakes, had scarcely two specimens alike, and many of them closely resembled other and altogether distinct species.[17] _The Variability of Insects_.
Among Insects there is a large amount of variation, though very few entomologists devote themselves to its investigation.


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