All are studious.
S.Matthew, on the upper left as one stands with one's back to the Titian, has an open-air study, and he makes notes as he reads.
His eagle is in attendance.
S.Mark, with his lion at ease under his chair, has also his open-air desk, and as he reads he thinks.
S. John is indoors, reading intently, with a box full of books to fall back on, and a little angel peeping at him from behind his chair.