We can thus understand the concentration of related species within the same areas; and how it is that under different latitudes, for instance, in South America, the inhabitants of the plains and mountains, of the forests, marshes, and deserts, are linked together in so mysterious a manner, and are likewise linked to the extinct beings which formerly inhabited the same continent.

Bearing in mind that the mutual relation of organism to organism is of the highest importance, we can see why two areas, having nearly the same physical conditions, should often be inhabited by very different forms of life; for according to the length of time which has elapsed since the colonists entered one of the regions, or both; according to the nature of the communication which allowed certain forms and not others to enter, either in greater or lesser numbers; according or not as those which entered happened to come into more or less direct competition with each other and with the aborigines; and according as the immigrants were capable of varying more or less rapidly, there would ensue in the to or more regions, independently of their physical conditions, infinitely diversified conditions of life; there would be an almost endless amount of organic action and reaction, and we should find some groups of beings greatly, and some only slightly modified; some developed in great force, some existing in scanty numbers--and this we do find in the several great geographical provinces of the world.
On these same principles we can understand, as I have endeavoured to show, why oceanic islands should have few inhabitants, but that of these, a large proportion should be endemic or peculiar; and why, in relation to the means of migration, one group of beings should have all its species peculiar, and another group, even within the same class, should have all its species the same with those in an adjoining quarter of the world.

We can see why whole groups of organisms, as batrachians and terrestrial mammals, should be absent from oceanic islands, whilst the most isolated islands should possess their own peculiar species of aerial mammals or bats.

We can see why, in islands, there should be some relation between the presence of mammals, in a more or less modified condition, and the depth of the sea between such islands and the mainland.

We can clearly see why all the inhabitants of an archipelago, though specifically distinct on the several islets, should be closely related to each other, and should likewise be related, but less closely, to those of the nearest continent, or other source whence immigrants might have been derived.